So, assuming the pump is ok and having good standby pressure of atleast 300psi which I think it does since you got the correct pilot pressure you should look at the load sense relief valve, located left rear of valvebody.
There's a new version that has a round body with hex shape only on the end that's supposed to be more reliable.
Then there's the load sense duplication valve located just below the ls relief valve, check the seals on that one.
If these looks ok and moves freely the next thing to check is the margin relief valve located front and center of the valve body.